Biome 4

Navigate Returnal's Biome 4, the Echoing Ruins. Learn enemy weaknesses, boss strategies for Hyperion, and how to find the Echoing Keys.

Biome 4 Enemies:

• Echoing Turret: Fires yellow lasers with a charge-up time. Destroy them before they fire or time your dash just before activation.
• Gyropod: Fires a quick stream of machinegun fire. A well-timed dash can evade this, but swarms can be tricky. They tend to get close.
• Proto-Strixera: Upgraded Strixera with a triple-stacked, fast-moving red ring attack that expands outwards. Dashing through it is recommended; jumping can be difficult on uneven terrain.
• Strixera Matriarch: Rare enemy. Has homing attacks, bullet spam, and a tricky purple laser attack that starts as four prongs and connects at your location. Dodge left or right as the beam is about to connect. It has a red weakpoint that exposes after taking enough damage; focus fire on it.
• Proto-Titanops: Similar to regular Titanops but with a new attack producing two purple rings that must be jumped or outrun. They are quicker and more persistent.
• Malformed Proto-Titanops: Another Titanops variant with more purple laser attacks.
• Trichozoid: If they have a red barrier, melee them to break it, then follow up with a few shots. If no barrier, attack from a distance; they are slow and don't move much.

Boss Battle: Hyperion

Hyperion is a challenging boss, known for his tankiness. Always move and strafe. He spends much of the fight immobile at his organ, making him an easy target, but he becomes very durable.

• Phase 1: Hyperion stays at the organ. Focus on dealing damage quickly. He will eventually use attacks from the organ itself. Stay at mid-range, strafe left or right, and change direction when you reach the arena's edge. Most attacks are too slow to hit a constantly moving player.
• Phase 1 Attacks:
  • Two Volleys of Blue Homing Bullets: Move and dash left or right to avoid.
  • Red Ground Ring: A red energy ring projects outwards from the ground.